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HYMENOPTERA, Apidae (Apoidea) -- <Images>
& <Juveniles> Description &
Statistics
Apidae includes the honeybee and
some of the socialized bees, but the genus Psithyrus parasitizes the bumblebees (Clausen
1940/62). This family includes
long-tongued bees without pygidial or basitibial plates (Finnamore &
Michener 1993). There is no scopa in
queens of social species and in the parasitic and robber genera, The honeybees always
produce a wax comb with hexagonal cells. The cells serve for larval rearing
sites and honey storage. Honey is
formed in the stomach from nectar through the action of enzymes. It is regurgitated into the storage cells. The developing brood is fed with
pollen. The queen is responsible for
the production of eggs. She produces
a "queen substance" that suppresses the development of other
females in the colony. A marked division of labor occurs
in the colony. The drones
exist solely for the purpose of mating with the queen. There is a continuous cycle in a colony,
and a division takes place when a second queen is produced. The old queen then leads a part of the old
colony away to a new site in a swarm.
Three principal stimuli to the production of new queens are, (1) when
an overabundance of individuals occurs in the hive, (2) an old queen dies and
(3) when there is a shortage of food.
The latter case stimulates swarming to form new colonies. Apiculture regularly includes
artificial insemination. The genetic
configureation of a queen is 2X, a worker 2X and a drone 1X. Drones are produced from unfertilized
eggs. Honeybees are of great economic
importance in that they are widely deployed for the pollination of both
orchard and field crops (Please see ). Bee venom has been
used in therapy and royal jelly has been touted for rather doubtful
rejuvenation properties. The family includes about. 1,000
species including all the highly social bees as well as some solitary and
primitively social forms. There were
47 species known in North America as of 2000. Principal subfamilies are:
Euglossinae, Bombinae, Meliponinae and Apinae. One species, A. mellifera
L., has been transported worldwide for pollination and honey production. The earlier range was from northern Europe
to southern Africa. The other species
are found in southern and eastern Asia.
Colonies are perennial and they swarm. The old queen departs the nest with many workers. Queens and workers differ in
appearance. Colonies generally have
thousands of bees and in the wild occur in hollow trees, rock or soil
cavities. In southern Asia they have
been found on exposed combs of cells hanging from tree branches or ledges
(Finnamore & Michener 1993). This
is a large family of bees, comprising the common honey bees, stingless bees
(which are also cultured for honey), carpenter bees, orchid bees, cuckoo
bees, bumblebees, and various other less well-known groups. The family Apidae
presently includes all the genera that were previously classified in the
families Anthophoridae and Ctenoplectridae, and most of these are solitary
species, though a few are also cleptoparasites. The four groups that were
subfamilies in the old family Apidae are presently ranked as tribes within
the subfamily Apinae. This trend has been taken to its extreme in a few
recent classifications that place all the existing bee families together
under the name "Apidae" (or, alternatively, the non-Linnaean clade
"Anthophila"), but this is not a widely-accepted practice.
The subfamily Apinae contains a diversity of lineages, the majority of
which are solitary, and whose nests are simple burrows in the soil. However, honey
bees, stingless bees, and bumblebees are colonial (eusocial), though they are
sometimes believed to have each developed this independently, and show
notable differences in such things as communication between workers and
methods of nest construction. Xylocopines (the subfamily which includes
carpenter bees) are mostly solitary, though they tend to be gregarious, and
some lineages such as the Allodapini contain eusocial species; most members
of this subfamily make nests in plant stems or wood. The nomadines are all
cleptoparasites in the nests of other bees. Other key
references are Maa (1953), Schwarz (1939, 1948) and Michener (1990). References: Please refer
to <Link 2>, [Additional
references may be found at: Link
1 ] |