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Table 1. Genera of Coleoptera which predominate in each intertidal zone

in southern California

Rocky headlands and reefs* ---------------------------------- (a) Area of-red algae down to mean low water Diaulota (b) Area of high tides Thalassotrechus* Diglotta* Bryothinusa* Diaulota* (also Aegialites. Amblopusa*, Endeodes Liparocephalus* and Ochthebius in central California) (c) Area of spray Bryothinusa* Diaulota* Thalassotrechus

Sandy beaches ---------------------------------- Area of fresh seaweed Cafius* Cicindela Dyschirius Thinopinus* Area of decaying seaweed Aleochara* Anthicus Bledius* Bryobiota* Cafius* Cercyon Diglotta* Emphyastes Hadrotes* Hypocaccus Motschulkium Neopachylopus Phaleria Phycocoetes Pontamalota* Tarphiota* Thinusa* Area of dry seaweed Amblyoderus Anthicus Apsena Catorama Endeodes Phaleria Phyconomus

Mudflats ----------------------------------- Area of daily tides Berosus Enochrus Tropisternis (All in the water) Area of sea-lettuce Actinidium Bembidion* Carpelimus* Cicindela Myllaena* Ochthebius Tachys Thinobius* Throscinus Area of Salicornia Bledius* Carpelimus* Thinobius*

* Staphyhinidae.

continue their activities in seawater at a reduced rate of metabolism (Meyerdirk, 1969). All of the species, which inhabit the reef area, are submarine as they are normally submerged in seawater part of the time. One species, Diaulota harteri Moore, is found as deep as mean low water where it would sometimes be submerged for as long as 20 hours a day. These species spend most of their time in crevices in the rocks or in empty barnacle shells, etc. They sometimes come out and wander about at low tide.

It is questionable if any of the typical species of the sandy beach could be called