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          Table 1.  Genera of Coleoptera which predominate in each intertidal zone

                 in southern California

 

Rocky headlands and reefs*

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(a) Area of-red algae down

       to mean low water

 

       Diaulota

 

 

 

(b) Area of high tides

 

    Thalassotrechus*

    Diglotta*

    Bryothinusa*

    Diaulota*

    (also Aegialites.

      Amblopusa*, Endeodes

      Liparocephalus* and

      Ochthebius in central

      California)

 

 

 

 

 

 

 

 

 

(c)  Area of spray

 

    Bryothinusa*

    Diaulota*

    Thalassotrechus

 

 

 

 

Sandy beaches

 

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Area of fresh seaweed

 

    Cafius*

    Cicindela

    Dyschirius

    Thinopinus*

 

Area of decaying seaweed

 

    Aleochara*

    Anthicus

    Bledius*

    Bryobiota*

    Cafius*

    Cercyon

    Diglotta*

    Emphyastes

    Hadrotes*

    Hypocaccus

    Motschulkium

    Neopachylopus

    Phaleria

    Phycocoetes

    Pontamalota*

    Tarphiota*

    Thinusa*

 

Area of dry seaweed

 

    Amblyoderus

    Anthicus

    Apsena

    Catorama

    Endeodes

    Phaleria

    Phyconomus

Mudflats

 

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Area of daily tides

 

    Berosus

    Enochrus

    Tropisternis

   (All in the water)

 

Area of sea-lettuce

 

    Actinidium

    Bembidion*

    Carpelimus*

    Cicindela

    Myllaena*

    Ochthebius

    Tachys

    Thinobius*

    Throscinus

 

 

 

 

 

 

 

 

 

Area of Salicornia

 

    Bledius*

    Carpelimus*

    Thinobius*

 

 

 

 

* Staphyhinidae.

 

continue their activities in seawater at a reduced rate of metabolism (Meyerdirk, 1969). All of the species, which inhabit the reef area, are submarine as they are normally submerged in seawater part of the time. One species, Diaulota harteri Moore, is found as deep as mean low water where it would sometimes be submerged for as long as 20 hours a day. These species spend most of their time in crevices in the rocks or in empty barnacle shells, etc. They sometimes come out and wander about at low tide.

 

          It is questionable if any of the typical species of the sandy beach could be called