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18. Mandibles reduced (a1), not capable of meeting medially. Basal metatarsal segment (b1) shorter than 2nd segment, subequal or shorter than mesotibial spur. Flagellum usually 3-segmented in females (c1) (always 4-segmented in males). Postmarginal vein subequal or slightly longer than stigmal vein (at most 1.7x stigmal vein length).
18'. Mandibles not strongly reduced (aa1), capable of meeting medially. Basal metatarsal segment equal or longer than second metatarsal segment and metatibial spur. Flagellum often 4-segmented in females (3-segmented in male Colpoclypeus). Postmarginal vein (dd1) often more than 1.7x stigmal vein length.
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Image credits: a1, aa1: Schauff, et al. (1997). c1: Askew (1968). dd1: Boucek (1988)