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Key to the Chalcidoidea of New Zealand
(derived
fr. Noyes & Valentine, 1989 & Valentine, 1971 by E. F. Legner)
[This key is in a form commonly used in North America. If the statement is true, proceed to
the
designated couplet, whereas if it is false, go to "b" portion of the
couplet. Numbers in
parentheses refer to previous couplet read].
Please refer to <Guide> for illustrations (Powerful connections use <Guide-2>)]
Keys to New Zealand Chalcidoidea:
1a. Abdominal petiole long, with 2 segments
(Fig. A); fore wing (when present) pedunculate, varying in shape but
commonly spatulate with a long marginal fringe (Fig. A), disc
reticulately alveolate (Fig. A), and venation
reduced and indistinct; pronotum reaches tegulae, without an
intervening prepectus; minute. <0.75 mm.
long ..........................................................................Mymarommatidae [Noyes & Valentine (1989) placed
this family in the Mymarommatoidea].
1b. Abdominal petiole with 1 segment, often
indistinct or somewhat completely invisible; fore wing not as
previous, if pedunculate then not reticulate; pronotum does not
reach tegulae, separated from them by a
prepectus (Fig. A); normally longer than 0.75 mm., but sometimes
much shorter........(Chalcidoidea)......2
2a
(1). Tarsi with 34 segments; antennae
short, the funicle with no more than 2 segments (Figs. A-E); minute to
very small insects, not more than 1.2 mm. long
..........................................................Trichogrammatidae
2b. Tarsi with 4-5 segments; minute to
moderate-sized insects varying from 0.2 mm. to more than 10 mm. long,
mostly exceeding 1.2
mm..............................................................................................................................3
3a
(2). Antennal toruli situated much
closer to eyes than to each other; frons with a straight, transverse suture a
little above toruli which connects with vertical sutures adjacent
to each orbit, thus forming an H
(Figs. A-
B); the winged species almost always with membrane of hind wing not
extending to base, thus giving hind
wing a stalked appearance (Fig. C) [exception = Anagroidea
spp. (Fig. D)]..............................Mymaridae
3b. Antennal toruli situated as close to or
closer to each other than to eyes or very nearly so; frons sometimes with
a transverse suture, which may be straight or V-shaped, but never
with vertical sutures running adjacent to
inner orbits; the winged species with membrane of hind wing always
extending to base............................4
4a
(3). Either winged, with stigmal vein
of fore wing long and forming an angle of ca. 90 deg. with marginal vein
(Figs. A-B), or completely apterous (males only), with eyes minute,
less than 1/2 as long as distance
between them (Fig. C).
Males with fore and hind femora very stout, contrasting with relatively
slender
middle femora (Fig. C)
[associated with fig
fruit].........................................................................Agaonidae
4b. Without previous combination of
characters......................................................................................................5
5a
(4). Hind leg with femur swollen and
tibia curved; body black, without metallic hue......................Chalcididae
5b. Either hind leg with femur not swollen and
tibia straight, or head and thorax metallic green...........................6
6a
(5). Hind coxa elongated, at least ca.
2X as long as fore coxa (Fig. A); fore wings fully developed, with stigmal
vein short and uncus hardly separated from the well developed
postmarginal vein (Figs. A-C); ovipositor
often well exserted
.......................................................................................................................Torymidae
6b. Hind coxa not so enlarged, not or hardly
longer than fore coxa; fore wing, if fully developed, usually with
stigmal vein longer and uncus well separated from postmarginal
vein, or postmarginal vein absent;
ovipositor normally hardly exserted .............................................................................................................7
7a
(6). Antennae with very long,
unsegmented clava and funicle composed of 2-4 indistinct, strongly transverse
segments (Figs. A-B); body shining black or yellow; gaster sessile;
axillae not distinctly marked off from
scutellum, the two together forming a strongly transverse band ca.
3X as broad as long; propodeum with a
large, shiny, central triangular area (Fig.
C)............................................................................Signiphoridae
7b. Antennae not as previous; occasionally
clava long and unsegmented, but then funicle composed of only 2
strongly transverse segments, and body metallic green; scutellum
shield-shaped, almost as long as broad or
slightly transverse, usually with axillae distinctly marked off;
propodeum without a distinct triangular
central
area....................................................................................................................................................8
8a
(7). All tarsi with 4
segments..............................................................................................................................9
8b. At least hind tarsi have 5
segments.................................................................................................................13
9a
(8). Fore wing marginal vein
indistinct, somewhat punctiform (Fig.
A)............................................Encyrtidae
9b. Fore wing marginal vein distinct, several
times longer than
broad................................................................10
10a
(9). Antennae with 14 segments; funicle
and clava each composed of 6 segments (Fig. A)..............Rotoitidae
10b. Antennae with not more than 12 segments;
funicle composed of not more than 5 segments, and clava never
with more than 3
segments........................................................................................................................11
11a
(10). Hind leg with coxa strongly
expanded, disc-shaped (Fig. A) and tibia on outer surface with coarse, dark
bristles arranged in longitudinal rows or diamond-shaped patterns;
gaster in cross-section somewhat
triangular......................................................................................................................................Elasmidae
11b. Hind leg with coxa subcylindrical, not
compressed, and hind tibia without darker bristles arranged in a
conspicuous pattern; gaster not triangular in
cross-section.......................................................................12
12a
(11). Gaster distinctly constricted
at junction with propodeum; fore wing in winged forms with postmarginal
and stigmal veins often long and distinct; body almost always at
least partly metallic; notaular lines, if
complete, almost always curved
................................................................................................Eulophidae
12b. Gaster at base about as broad as propodeum,
not distinctly constricted; fore wing in macropterous forms with
postmarginal vein absent, or almost so, and stigmal vein very short
(Figs. A-E); body not metallic, usually
brown or black; notaular lines complete, straight (Figs. C-D,
F)................................................Aphelinidae
13a
(8). Either mesopleuron undivided,
relatively large and shield-shaped (Figs. A-C) or gaster broadly sessile
(Figs. D-G); middle tibia usually with a strong apical spur (Figs.
B, D, F, H, I-K)..................................14
13b. Mesopleuron divided into mesepisternum and
mesepimeron (Fig. A), the 2 parts often with distinctly
different sculpture; gaster never broadly sessile, at least with a
distinct constriction at junction with
propodeum, often petiolate; middle tibia with spur of normal
proportions (Figs. B-G)...........................16
14a
(13). Thorax in profile with middle
coxa inserted about level with middle of mesopleural shield (Fig. A) or
even slightly anterior to this; fore wing, if fully developed, with
marginal vein short, usually not more than
3-4X as long as broad
.................................................................................................................Encyrtidae
14b. Thorax in profile with middle coxa
inserted about level with posterior margin of mesopleural shield (Figs. A-
C); fore wing, if fully developed, with marginal vein always at
least 6-7X as long as broad.....................15
15a (14). Antennae with flagellum having not more
than 7 segments; gaster sessile, broadly attached to
propodeum (Figs. A-F); mesoscutum at least slightly convex, with
notauli always present and straight
(Figs. A-B, G); length not more than 1.5 mm ...........................................................................Aphelinidae
15b. Antennae with flagellum having 8-9
segments; gaster distinctly constricted at junction with propodeum, or
petiolate (Figs. A-B); mesoscutum either impressed or convex, with
notauli very inconspicuous; length
almost always >1.5
mm.............................................................................................................Eupelmidae
16a
(13). Pronotal collar large,
subrectangular, at least about 2/3rds as long as mesoscutum (Figs. 58, 64);
antennae with not more than 6 funicle segments; head and dorsum of
thorax with numerous conspicuous,
hairy punctures which often give rise to very coarse sculpture;
gena sharply margined
posteriorly...............................................................................................................................Eurytomidae
16b. Pronotal collar not large and
subrectangular, shorter than 1/2 length of mesoscutum, or if longer then
antennae with 7 funicle segments, or sculpture of head and thorax
shallow, or gena not with a sharp
edge.............................................................................................................................................................17
17a (16). Fore wing with apex of stigmal vein much
enlarged, deeper than long, and apex of uncus very close to
postmarginal vein (Fig. A); pronotum elongated, subconical; female
with ovipositor strongly
exserted.........................................................................................................................................Torymidae
17b. Fore wing with apex of stigmal vein not or
hardly enlarged, or if as above then pronotum transverse and not
well developed; female rarely with ovipositor strongly
exserted...............................................................18
18a
(17). Wings fully developed; fore
wing marginal vein at least 3.5X as long as stigmal vein (Fig. A); gastral
petiole at least 1.5X as long as broad; antennae inserted above
lowest eye margins, without a crest or
tubercle between
them..............................................................................................................Perilampidae
18b. If wings fully developed, then fore wing
marginal vein less than 3.5X as long as stigmal vein, or if relatively
longer then either gastral petiole not longer than broad, or
antenna inserted well below lowest margin of
eye, or a sharp crest or tubercle present between antennal
toruli...............................................................19
19a
(18). Mesopleuron divided by a very
weak, inconspicuous depression; notaular grooves complete; fore wing
marginal vein always more than 2X as long as stigmal vein; antenna
with 7 funicle segments and a single
small anellus; scutellum conspicuously hairy; males only (Fig.
A)............................................Eupelmidae
19b. Mesopleuron distinctly divided into
episternum and epimeron; notaular grooves sometimes incomplete;
marginal vein mostly less than 2X as long as stigmal vein; antennae
usually with 6 or fewer funicle
segments, often with as many as 3 anelli; scutellum usually not
conspicuously hairy; males and females
.................................................................................................................................................Pteromalidae
REFERENCES:
Noyes, J.
S. & E. W. Valentine. 1989. Chalcidoidea (Insecta: Hymenoptera)-- Introduction,
and Review of Genera
in Smaller Families. Fauna of New Zealand No. 18. DSIR Publ., Wellington, New Zealand. 58 p.
Valentine,
E. W. 1971. Entomology of the Aucklands and other islands south of New
Zealand: Hymenoptera
Mymaridae. Pacific Insects Monog. 27:
327-33.